ASIAN RED PAPER WASP (Ropalidia marginata) ©Rundstedt-Rovillos / Solent
Primitively eusocial societies are typically headed by behaviourally aggressive queens, who use aggression to suppress worker reproduction. The queen in R. marginata, however, is a “docile sitter” who does not use physical aggression to maintain her reproductive monopoly in the colony.
The queen is also not responsible for maintaining worker activity in her colony, as in similar species. However, the queen in R. marginata is able to maintain complete reproductive monopoly in the colony. It has been argued for several years that the R. marginata queen uses a pheromone to signal her presence and fecundity to her workers, and this signal is perceived by the workers who refrain from reproducing; however, it has been shown that any such pheromone is non-volatile.
Fact Source: http://en.wikipedia.org/wiki/Ropalidia_marginata
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